The film begins at Flemner Air Base 20 years in the past. A pilot named Leland "Buzz" Harley (Bill Irwin) loses control of his plane and ejects, leaving his co-pilot Dominic "Mailman" Farnham (Ryan Stiles) to crash; although Mailman survives, he's mistaken for a deer owing to the branches stuck to his helmet and is shot by a hunter. Topper Harley (Charlie Sheen) wakes up from a nightmare he's having about the event when Lt. Commander Block (Kevin Dunn) asks him to return to active duty as a pilot in the U.S. Navy, to help on a new top secret mission: Operation Sleepy Weasel, commanded by the incompetent and oblivious Admiral Benson (Lloyd Bridges). Harley starts to show some psychological problems, especially when his father is mentioned. His therapist, Ramada (Valeria Golino), tries to keep Topper from flying, but she relents, and also starts to build a budding romance with Topper. Meanwhile, Topper gets into a rivalry with another fighter pilot, Kent Gregory (Cary Elwes), a former lover of Ramada and son of "Mailman" Farnham who blames Buzz Harley for his father's death, and believes Topper cannot handle combat pressure.